Fruit Ripening

Ripening is programmed senescence — a coordinated enzymatic self-destruction that converts a seed-protecting structure into a seed-dispersing reward. Understanding the biochemistry of ripening is the single most useful piece of knowledge for buying, storing, and cooking fruit, because it determines whether a fruit can improve after harvest or is locked in at the moment it was picked.

Four stages of fruit development

Fruits develop through fertilization and hormone induction, cell multiplication (brief), cell expansion (the major growth phase, where storage cells fill with water, sugars, defensive compounds, and pre-positioned enzyme systems), and finally ripening itself. During the expansion phase, melon fruits can grow 80 cc daily; watermelon cells reach visible millimeter scale.